"The hunting of vertebrates can also affect invertebrate populations. For
example, a reduction in the amount of dung directly stresses the populations
of dung beetles."
Biotropica 39, 328; 289 (2007).
I never thought of "dung beetle" as a high-stress job.
Others, however, reinforce this notion:
Well not until there's no shit to dung around in.
-Veronique Chez Sheep
Would YOU want to do it?
-Bill in Vancouver
Most dung beetles quit their jobs after a couple of years.
So if your dung beetles can't meet their quota, what kind of job do you
threaten them with to motivate them?
Use scarab tactics